The Gloucester midfield specialist has been drafted in by coach Andy Robinson to provide the cutting edge and dash to an England team who have scored only three tries in two games against the Wallabies and the All Blacks.
Simpson-Daniel has been in electric form for Gloucester from the number 13 position, combining well with youngster Anthony Allen and Test star Mike Tindall after switching from the wing.
He comes into the side for Jamie Noon and told www.gloucesterrugby.co.uk "It's a great surprise to be named again but I had been in contact with Andy Robinson before the Leicester game because he wanted to know how things were going for me.
"I am delighted to be back involved with the England squad, no matter what the circumstances, and now the chance is there for me, I am desperate to try and take it.
"I suppose I am under a bit of pressure but I just have to try and do the best I can, make a contribution to the team and give it everything I have got.
"I am really looking forward to just having that chance again - whether it came on the wing or in the centre. I am certainly getting more and more confident when I play for Gloucester in the centre and I want to take that forward now with England.
"When I missed out on the squad originally, all I could do was come back to the club and try and play as well as I could so if the opportunity came my way, I was in a position to take it."
Simpson-Daniel will be partnered in the centre by his Kingsholm team-mate Tindall and the duo have been paired together only twice for Gloucester this season against Bayonne and Bucuresti in the European Challenge Cup.
Gloucester number eight James Forrester has also included and will sit on the bench. Phil Vickery, who started at tighthead against both Australia and the All Blacks, should be available for Gloucester's Guinness Premiership match against Saracens.
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